Ryan P. Walter

Contact Information

Office: MH-689A

Phone: 657-278-4812


Ryan P. Walter

associate Professor


PhD, University of Windsor

MS, State University of New York at Brockport

BS, State University of New York at Brockport

Research Areas

Molecular ecology, hybridization, phylogeography, population connectivity

Courses Regularly Taught

Evolutionary Genetics, Genetics, Elements of Biology


Taboun Z, Walter RP, Ovenden JR, Heath DD (2021) Spatial and temporal genetic variation in an exploited reef fish: the effects of exploitation on cohort genetic structure. Evolutionary Applications, 14: 1286-1300. 

Swintek MA, Walter RP (2021) Development and characterization of thirteen novel microsatellite markers for use in Greenland sharks (Somniosus microcephalus), with cross-amplification in Pacific sleeper sharks (Somniosus pacificus). BMC Research Notes, 14:28. 

Roy, D, Lehnert SJ, Venney CJ, Walter R, Heath DD (2021) NGS-µsat: Bioinformatics framework supporting high throughput microsatellite genotyping from next generation sequencing platforms. Conservation Genetics Resources, 13:161-173. 

Lowe A, Summers AP, Walter RP, Walker S, Paig-Tran EWM (2020) Scale performance and composition in a small Amazonian armored catfish, Corydoras trilineatus. Acta Biomaterialia, S1742-7061(20)30699-1. 

Carrillo Munoz N, Dickson KA, Walter RP (2020) The mitogenome of the California grunion (Leuresthes tenuis). Mitochondrial DNA Part B. 5:1138–1140 doi: 10.1080/23802359.2020.1730254

Edwards JE, Broell F, Bushnell PG, Campana SE, Christiansen JS, Devine BM, Gallant JJ, Grant SM, Hedges KJ, Hiltz E, MacNeil MA, McMeans BC, Nielsen J, Præbel K, Skomal GB, Steffensen JF, Walter RP, Watanabe YY, VanderZwaag DL, Hussey NE (2019) Advancing our understanding of long-lived species: A case study on the Greenland shark. Frontiers in Marine Science. 6:87 doi: 10.3389/fmars.2019.00087.

Graham BA, Heath DD, Walter RP, Mark MM, Mennill DJ (2018) Parallel evolutionary forces influence the evolution of male and female songs in a tropical songbird. Journal of Evolutionary Biology. 31: 979-994. 

Graham BA, Heath DD, Walter RP, Mennill DJ (2018) Immigrant song: males and females learn songs post-dispersal in a tropical bird. Behavioural Ecology 29:711-723.

Lisi PJ, Childress ES, Gagne RB, Hain EF, Lamphere BA, Walter RP, Hogan JD, Gilliam JF, Blum MJ, McIntyre PB (2018) Overcoming urban stream syndrome: trophic flexibility confers resilience among Hawaiian stream fishes. Freshwater Biology. DOI: 10.1111/fwb.13091.

Balasingham KD, Walter RP, Mandrak NE Heath DD (2018) Environmental DNA detection of rare and invasive fish species in two Great Lakes tributaries. Molecular Ecology 27:112-127. 

Kessel ST, Alhasan N, Yurkowski DJ, Walter RP, Klaus R, Chekchak T, Hill G, Hussey NE (2017) Conservation of reef manta rays (Manta alfredi) in a UNESCO World Heritage Site: Large-scale island development or sustainable tourism? PLoS ONE 12(10):e0185419.

Walter RP, Roy D, Hussey NE, Stelbrink B, Kovacs KM, Lydersen C, McMeans BC, Svavarsson J, Kessel ST, Porsmoguer SB, Wildes S, Tribuzio CA, Campana SE, Petersen SD, Grubbs RD, Heath DD, Hedges KJ, Fisk AT (2017) Origins of the Greenland shark (Somniosus microcephalus): impacts of ice-olation and introgression. Ecology and Evolution 7:8113–8125. 

Moody KN, Gagne RB, Heim-Ballew H, Alda F, Hain EF, Lisi P, Walter RP, Higashi GR, Hogan JD, McIntyre PB, Gilliam JF, Blum MJ (2017) Invasion hotspots and ecological saturation of streams across the Hawaiian archipelago. Cybium 41:127-156.

Hinojosa-Alvarez S, Walter RP, Diaz-Jaimes P, Galván-Magaña F, Paig-Tran EM (2016) A potential third Manta Ray species near the Yucatán Peninsula? Evidence for a recently diverged and novel genetic Manta group from the Gulf of Mexico. PeerJ 4:e2586 

Balasingham KD, Walter RP, Heath DD (2016) Residual eDNA detection sensitivity assessed by quantitative real-time PCR in a river ecosystem. Molecular Ecology Resources. DOI: 10.1111/1755-0998.12598

Alda F, Gagne RB, Walter RP, Hogan JD, Moody K, Zink F, McIntyre PB, Gilliam JF, Blum MJ (2016) Colonization and demographic expansion of amphidromous stream fauna across the Hawaiian Archipelago. Journal of Evolutionary Biology 29:2054-2069.

Roy D, Lucek K, Walter RP, Seehausen O (2015) Hybrid ‘superswarm’ leads to rapid divergence and establishment of populations during a biological invasion. Molecular Ecology 24:5394-5411.

Glass WR, Walter RP, Heath DD, Mandrak NE, Corkum LD (2015) Genetic structure and diversity of Spotted Gar (Lepisosteus oculatus) at its northern range edge: Implications for conservation. Conservation Genetics 16:889-899.

Ginson R, Walter RP, Mandrak NE, Beneteau CL, Heath DD (2015) Hierarchical analysis of genetic structure in the habitat-specialist Eastern Sand Darter (Ammocrypta pellucida). Ecology and Evolution 5:695-708.

Hussey NE, Cosandey-Godin A, Walter RP, Hedges KJ, VanGerwen-Toyne M, Barkley AN, Kessel ST, Fisk AT (2015) Juvenile Greenland sharks Somniosus microcephalus (Bloch & Schneider, 1801) in the Canadian Arctic. Polar Biology 38:493-504.

Walter RP, Kessel ST, Alhasan N, Fisk AT, Heath DD, Chekchak T, Klaus R, Younis M, Hill G, Jones B, Braun CD, Berumen ML, DiBattista JD, Priest MA, Hussey NE (2014) First record of living Manta alfredi × Manta birostris hybrid. Marine Biodiversity 44:1-2.

Smit J, Heath DD, Walter RP (2014) A molecular genetic lab to generate inclusive and exclusive forensic evidence: two suspects, a victim and a bloodstained t-shirt. Biochemistry and Molecular Biology Education 42:230-236.

Walter RP, Cena CJ, Morgan GE, Heath DD (2012) Historical and anthropogenic factors affecting the population genetic structure of Ontario's inland lake populations of walleye (Sander vitreus). Journal of Heredity 103:831-841.

Herder F, Schliewen UK, Geiger MF, Hadiaty RK, Gray SM, McKinnon JS, Walter RP, Pfaender J (2012) Alien invasion in Wallace's Dreampond: records of the hybridogenic "flowerhorn" cichlid in Lake Matano, with an annotated checklist of fish species introduced to the Malili Lakes system in Sulawesi. Aquatic Invasions 7:521-535.

Walter RP, Hogan JD, Blum MJ, Gagne RB, Hain EF, Gilliam JF, McIntyre PB (2012) Climate change and conservation of endemic amphidromous fishes in Hawaiian streams. Endangered Species Research 16:261-272. 

Lindstrom DP, Blum MJ, Walter RP, Gagne RB, Gilliam JF (2012)  Molecular and morphological evidence of distinct evolutionary lineages of Awaous guamensis in Hawai'i and Guam. Copeia 2012:294-301.

Beneteau CL, Walter RP, Mandrak NE, Heath DD (2012) Range expansion by invasion: genetic characterization of invasion of the Greenside Darter (Etheostoma blennioides) at the northern edge of its distribution. Biological Invasions 14:191-121.

Walter RP, Hogan JD, Haffner GD, Heath DD (2011) Genetic structure and connectivity among lake populations of threatened Paratherina sailfin silversides from Sulawesi, Indonesia. Conservation Genetics 12:1387-1393. 

Walter RP, Blum MJ, Snider SB, Paterson IG, Benzten P, Lamphere BA, Gilliam JF (2011) Isolation and differentiation of Rivulus hartii across Trinidad and neighboring islands. Molecular Ecology 20:608-618.

Salas E, Molina H, Walter RP, Heath DD (2010) Local and regional genetic connectivity in a Caribbean coral reef fish. Marine Biology 157:437–445.